Theoretical morphology · 1966

Raup's Cube

Every coiled shell is one tube that grows as it turns. Three numbers decide how it looks.

Step 1 / 6

One tube

A shell is a single tube that grows as it turns. The amber ring is the opening, sweeping around an axis and getting bigger at a steady rate. Every whorl is a scaled copy of the one before: a logarithmic spiral. This one is a nautilus.

WWhorl expansion

How much the tube grows in one turn

DDistance from the axis

Inner edge ÷ outer edge of the tube

TTranslation

How far the tube slides down the axis

Morphospace

The cube

Each point is a shell. Drag across the map to set W and D, and the strip on the right for T. Below the amber line, whorls touch.

Regions are approximate, after Raup (1966). The model assumes a circular tube and steady growth, so real shells only sit near their points.

About the model

In the 1960s the paleontologist David Raup asked a computer to draw shells. He found that one rule covers nautiluses, ammonites, snails, clams and brachiopods: a tube that keeps its shape while it grows and turns around an axis. Three numbers set its form, so all possible shells fit in one cube, and real animals use only a small part of it.